diff --git a/crates/bevy_pbr/Cargo.toml b/crates/bevy_pbr/Cargo.toml
index bf568885e7fef..5031d7c91444f 100644
--- a/crates/bevy_pbr/Cargo.toml
+++ b/crates/bevy_pbr/Cargo.toml
@@ -37,6 +37,7 @@ bevy_asset = { path = "../bevy_asset", version = "0.16.0-dev" }
 bevy_color = { path = "../bevy_color", version = "0.16.0-dev" }
 bevy_core_pipeline = { path = "../bevy_core_pipeline", version = "0.16.0-dev" }
 bevy_derive = { path = "../bevy_derive", version = "0.16.0-dev" }
+bevy_diagnostic = { path = "../bevy_diagnostic", version = "0.16.0-dev" }
 bevy_ecs = { path = "../bevy_ecs", version = "0.16.0-dev" }
 bevy_image = { path = "../bevy_image", version = "0.16.0-dev" }
 bevy_math = { path = "../bevy_math", version = "0.16.0-dev" }
diff --git a/crates/bevy_pbr/src/meshlet/instance_manager.rs b/crates/bevy_pbr/src/meshlet/instance_manager.rs
index f615114d417a1..26f6432a1fdf7 100644
--- a/crates/bevy_pbr/src/meshlet/instance_manager.rs
+++ b/crates/bevy_pbr/src/meshlet/instance_manager.rs
@@ -125,7 +125,6 @@ impl InstanceManager {
             None,
             None,
             None,
-            None,
         );
 
         // Append instance data
diff --git a/crates/bevy_pbr/src/render/gpu_preprocess.rs b/crates/bevy_pbr/src/render/gpu_preprocess.rs
index 26559f9223dd5..7cd757a2e5c25 100644
--- a/crates/bevy_pbr/src/render/gpu_preprocess.rs
+++ b/crates/bevy_pbr/src/render/gpu_preprocess.rs
@@ -626,12 +626,10 @@ impl Node for EarlyGpuPreprocessNode {
                     continue;
                 };
 
-                // If we're drawing indirectly, make sure the mesh preprocessing
-                // shader has access to the view info it needs to do culling.
-                let mut dynamic_offsets: SmallVec<[u32; 1]> = smallvec![];
-                if !no_indirect_drawing {
-                    dynamic_offsets.push(view_uniform_offset.offset);
-                }
+                // Make sure the mesh preprocessing shader has access to the
+                // view info it needs to do culling and motion vector
+                // computation.
+                let dynamic_offsets = [view_uniform_offset.offset];
 
                 // Are we drawing directly or indirectly?
                 match *phase_bind_groups {
@@ -1317,6 +1315,11 @@ fn preprocess_direct_bind_group_layout_entries() -> DynamicBindGroupLayoutEntrie
     DynamicBindGroupLayoutEntries::new_with_indices(
         ShaderStages::COMPUTE,
         (
+            // `view`
+            (
+                0,
+                uniform_buffer::<ViewUniform>(/* has_dynamic_offset= */ true),
+            ),
             // `current_input`
             (3, storage_buffer_read_only::<MeshInputUniform>(false)),
             // `previous_input`
@@ -1361,11 +1364,6 @@ fn gpu_culling_bind_group_layout_entries() -> DynamicBindGroupLayoutEntries {
             8,
             storage_buffer_read_only::<MeshCullingData>(/* has_dynamic_offset= */ false),
         ),
-        // `view`
-        (
-            0,
-            uniform_buffer::<ViewUniform>(/* has_dynamic_offset= */ true),
-        ),
     ))
 }
 
@@ -1802,11 +1800,14 @@ impl<'a> PreprocessBindGroupBuilder<'a> {
         )
         .ok();
 
+        let view_uniforms_binding = self.view_uniforms.uniforms.binding()?;
+
         Some(PhasePreprocessBindGroups::Direct(
             self.render_device.create_bind_group(
                 "preprocess_direct_bind_group",
                 &self.pipelines.direct_preprocess.bind_group_layout,
                 &BindGroupEntries::with_indices((
+                    (0, view_uniforms_binding),
                     (3, self.current_input_buffer.as_entire_binding()),
                     (4, self.previous_input_buffer.as_entire_binding()),
                     (
diff --git a/crates/bevy_pbr/src/render/mesh.rs b/crates/bevy_pbr/src/render/mesh.rs
index a02b56e441a37..34d0debbb7c6b 100644
--- a/crates/bevy_pbr/src/render/mesh.rs
+++ b/crates/bevy_pbr/src/render/mesh.rs
@@ -1,4 +1,7 @@
-use crate::material_bind_groups::{MaterialBindGroupIndex, MaterialBindGroupSlot};
+use crate::{
+    material_bind_groups::{MaterialBindGroupIndex, MaterialBindGroupSlot},
+    skin::mark_meshes_as_changed_if_their_skins_changed,
+};
 use allocator::MeshAllocator;
 use bevy_asset::{load_internal_asset, AssetId, UntypedAssetId};
 use bevy_core_pipeline::{
@@ -8,6 +11,7 @@ use bevy_core_pipeline::{
     prepass::MotionVectorPrepass,
 };
 use bevy_derive::{Deref, DerefMut};
+use bevy_diagnostic::FrameCount;
 use bevy_ecs::{
     prelude::*,
     query::ROQueryItem,
@@ -163,7 +167,13 @@ impl Plugin for MeshRenderPlugin {
 
         app.add_systems(
             PostUpdate,
-            (no_automatic_skin_batching, no_automatic_morph_batching),
+            (
+                no_automatic_skin_batching,
+                no_automatic_morph_batching,
+                mark_meshes_as_changed_if_their_skins_changed
+                    .ambiguous_with_all()
+                    .after(mark_3d_meshes_as_changed_if_their_assets_changed),
+            ),
         )
         .add_plugins((
             BinnedRenderPhasePlugin::<Opaque3d, MeshPipeline>::default(),
@@ -536,17 +546,15 @@ pub struct MeshInputUniform {
     pub index_count: u32,
     /// The current skin index, or `u32::MAX` if there's no skin.
     pub current_skin_index: u32,
-    /// The previous skin index, or `u32::MAX` if there's no previous skin.
     pub previous_skin_index: u32,
     /// The material and lightmap indices, packed into 32 bits.
     ///
     /// Low 16 bits: index of the material inside the bind group data.
     /// High 16 bits: index of the lightmap in the binding array.
     pub material_and_lightmap_bind_group_slot: u32,
+    pub timestamp: u32,
     /// User supplied tag to identify this mesh instance.
     pub tag: u32,
-    /// Padding.
-    pub pad: u32,
 }
 
 /// Information about each mesh instance needed to cull it on GPU.
@@ -1104,6 +1112,7 @@ impl RenderMeshInstanceGpuBuilder {
         render_material_bindings: &RenderMaterialBindings,
         render_lightmaps: &RenderLightmaps,
         skin_indices: &SkinIndices,
+        timestamp: FrameCount,
     ) -> u32 {
         let (first_vertex_index, vertex_count) =
             match mesh_allocator.mesh_vertex_slice(&self.shared.mesh_asset_id) {
@@ -1122,7 +1131,6 @@ impl RenderMeshInstanceGpuBuilder {
                 ),
                 None => (false, 0, 0),
             };
-
         let current_skin_index = match skin_indices.current.get(&entity) {
             Some(skin_indices) => skin_indices.index(),
             None => u32::MAX,
@@ -1156,6 +1164,7 @@ impl RenderMeshInstanceGpuBuilder {
             lightmap_uv_rect: self.lightmap_uv_rect,
             flags: self.mesh_flags.bits(),
             previous_input_index: u32::MAX,
+            timestamp: timestamp.0,
             first_vertex_index,
             first_index_index,
             index_count: if mesh_is_indexed {
@@ -1169,7 +1178,6 @@ impl RenderMeshInstanceGpuBuilder {
                 self.shared.material_bindings_index.slot,
             ) | ((lightmap_slot as u32) << 16),
             tag: self.shared.tag,
-            pad: 0,
         };
 
         // Did the last frame contain this entity as well?
@@ -1595,6 +1603,7 @@ pub fn collect_meshes_for_gpu_building(
     render_material_bindings: Res<RenderMaterialBindings>,
     render_lightmaps: Res<RenderLightmaps>,
     skin_indices: Res<SkinIndices>,
+    frame_count: Res<FrameCount>,
 ) {
     let RenderMeshInstances::GpuBuilding(ref mut render_mesh_instances) =
         render_mesh_instances.into_inner()
@@ -1634,6 +1643,7 @@ pub fn collect_meshes_for_gpu_building(
                         &render_material_bindings,
                         &render_lightmaps,
                         &skin_indices,
+                        *frame_count,
                     );
                 }
 
@@ -1661,6 +1671,7 @@ pub fn collect_meshes_for_gpu_building(
                         &render_material_bindings,
                         &render_lightmaps,
                         &skin_indices,
+                        *frame_count,
                     );
                     mesh_culling_builder
                         .update(&mut mesh_culling_data_buffer, instance_data_index as usize);
@@ -1843,7 +1854,6 @@ impl GetBatchData for MeshPipeline {
 
         let current_skin_index = skin_indices.current.get(&main_entity).map(SkinIndex::index);
         let previous_skin_index = skin_indices.prev.get(&main_entity).map(SkinIndex::index);
-
         let material_bind_group_index = mesh_instance.material_bindings_index;
 
         Some((
diff --git a/crates/bevy_pbr/src/render/mesh_preprocess.wgsl b/crates/bevy_pbr/src/render/mesh_preprocess.wgsl
index 9bbe96e80d633..932bd58a1aaab 100644
--- a/crates/bevy_pbr/src/render/mesh_preprocess.wgsl
+++ b/crates/bevy_pbr/src/render/mesh_preprocess.wgsl
@@ -192,18 +192,28 @@ fn main(@builtin(global_invocation_id) global_invocation_id: vec3<u32>) {
     }
 #endif
 
+    // Was the mesh transform updated this frame?
+    let timestamp = current_input[input_index].timestamp;
+    let mesh_changed_this_frame = timestamp == view.frame_count;
+
     // Look up the previous model matrix.
     let previous_input_index = current_input[input_index].previous_input_index;
     var previous_world_from_local_affine_transpose: mat3x4<f32>;
-    if (previous_input_index == 0xffffffff) {
-        previous_world_from_local_affine_transpose = world_from_local_affine_transpose;
-    } else {
+    if (mesh_changed_this_frame && previous_input_index != 0xffffffffu) {
         previous_world_from_local_affine_transpose =
             previous_input[previous_input_index].world_from_local;
+    } else {
+        previous_world_from_local_affine_transpose = world_from_local_affine_transpose;
     }
     let previous_world_from_local =
         maths::affine3_to_square(previous_world_from_local_affine_transpose);
 
+    let previous_skin_index = select(
+        0xffffffffu,
+        current_input[input_index].previous_skin_index,
+        mesh_changed_this_frame
+    );
+
     // Occlusion cull if necessary. This is done by calculating the screen-space
     // axis-aligned bounding box (AABB) of the mesh and testing it against the
     // appropriate level of the depth pyramid (a.k.a. hierarchical Z-buffer). If
@@ -342,7 +352,7 @@ fn main(@builtin(global_invocation_id) global_invocation_id: vec3<u32>) {
     output[mesh_output_index].lightmap_uv_rect = current_input[input_index].lightmap_uv_rect;
     output[mesh_output_index].first_vertex_index = current_input[input_index].first_vertex_index;
     output[mesh_output_index].current_skin_index = current_input[input_index].current_skin_index;
-    output[mesh_output_index].previous_skin_index = current_input[input_index].previous_skin_index;
+    output[mesh_output_index].previous_skin_index = previous_skin_index;
     output[mesh_output_index].material_and_lightmap_bind_group_slot =
         current_input[input_index].material_and_lightmap_bind_group_slot;
     output[mesh_output_index].tag = current_input[input_index].tag;
diff --git a/crates/bevy_pbr/src/render/skin.rs b/crates/bevy_pbr/src/render/skin.rs
index c248821ccafd3..c3082caf1e490 100644
--- a/crates/bevy_pbr/src/render/skin.rs
+++ b/crates/bevy_pbr/src/render/skin.rs
@@ -4,6 +4,7 @@ use std::sync::OnceLock;
 use bevy_asset::Assets;
 use bevy_ecs::prelude::*;
 use bevy_math::Mat4;
+use bevy_render::mesh::Mesh3d;
 use bevy_render::sync_world::MainEntityHashMap;
 use bevy_render::{
     batching::NoAutomaticBatching,
@@ -25,7 +26,7 @@ use bevy_transform::prelude::GlobalTransform;
 pub const MAX_JOINTS: usize = 256;
 
 /// The location of the first joint matrix in the skin uniform buffer.
-#[derive(Component)]
+#[derive(Component, Clone, Copy)]
 pub struct SkinIndex {
     /// The byte offset of the first joint matrix.
     pub byte_offset: u32,
@@ -232,3 +233,18 @@ pub fn no_automatic_skin_batching(
         commands.entity(entity).try_insert(NoAutomaticBatching);
     }
 }
+
+pub fn mark_meshes_as_changed_if_their_skins_changed(
+    mut skinned_meshes_query: Query<(&mut Mesh3d, &SkinnedMesh)>,
+    changed_joints_query: Query<Entity, Changed<GlobalTransform>>,
+) {
+    for (mut mesh, skinned_mesh) in &mut skinned_meshes_query {
+        if skinned_mesh
+            .joints
+            .iter()
+            .any(|&joint| changed_joints_query.contains(joint))
+        {
+            mesh.set_changed();
+        }
+    }
+}
diff --git a/crates/bevy_pbr/src/render/skinning.wgsl b/crates/bevy_pbr/src/render/skinning.wgsl
index 92e977aeb1b92..038b293eed238 100644
--- a/crates/bevy_pbr/src/render/skinning.wgsl
+++ b/crates/bevy_pbr/src/render/skinning.wgsl
@@ -34,7 +34,7 @@ fn skin_model(
         + weights.z * joint_matrices.data[indexes.z]
         + weights.w * joint_matrices.data[indexes.w];
 #else   // SKINS_USE_UNIFORM_BUFFERS
-    let skin_index = mesh[instance_index].current_skin_index;
+    var skin_index = mesh[instance_index].current_skin_index;
     return weights.x * joint_matrices[skin_index + indexes.x]
         + weights.y * joint_matrices[skin_index + indexes.y]
         + weights.z * joint_matrices[skin_index + indexes.z]
@@ -57,11 +57,15 @@ fn skin_prev_model(
         + weights.z * prev_joint_matrices.data[indexes.z]
         + weights.w * prev_joint_matrices.data[indexes.w];
 #else   // SKINS_USE_UNIFORM_BUFFERS
-    let skin_index = mesh[instance_index].previous_skin_index;
-    return weights.x * prev_joint_matrices[skin_index + indexes.x]
-        + weights.y * prev_joint_matrices[skin_index + indexes.y]
-        + weights.z * prev_joint_matrices[skin_index + indexes.z]
-        + weights.w * prev_joint_matrices[skin_index + indexes.w];
+    let prev_skin_index = mesh[instance_index].previous_skin_index;
+    if (prev_skin_index == 0xffffffffu) {
+        return skin_model(indexed, weights, instance_index);
+    }
+
+    return weights.x * prev_joint_matrices[prev_skin_index + indexes.x]
+        + weights.y * prev_joint_matrices[prev_skin_index + indexes.y]
+        + weights.z * prev_joint_matrices[prev_skin_index + indexes.z]
+        + weights.w * prev_joint_matrices[prev_skin_index + indexes.w];
 #endif  // SKINS_USE_UNIFORM_BUFFERS
 }
 
diff --git a/crates/bevy_render/src/experimental/occlusion_culling/mesh_preprocess_types.wgsl b/crates/bevy_render/src/experimental/occlusion_culling/mesh_preprocess_types.wgsl
index 6d1199adb4282..cbf9b2dc46338 100644
--- a/crates/bevy_render/src/experimental/occlusion_culling/mesh_preprocess_types.wgsl
+++ b/crates/bevy_render/src/experimental/occlusion_culling/mesh_preprocess_types.wgsl
@@ -19,9 +19,9 @@ struct MeshInput {
     // Low 16 bits: index of the material inside the bind group data.
     // High 16 bits: index of the lightmap in the binding array.
     material_and_lightmap_bind_group_slot: u32,
+    timestamp: u32,
     // User supplied index to identify the mesh instance
     tag: u32,
-    pad: u32,
 }
 
 // The `wgpu` indirect parameters structure. This is a union of two structures.
diff --git a/crates/bevy_render/src/render_resource/buffer_vec.rs b/crates/bevy_render/src/render_resource/buffer_vec.rs
index 0feaa8487885b..32d06ab397578 100644
--- a/crates/bevy_render/src/render_resource/buffer_vec.rs
+++ b/crates/bevy_render/src/render_resource/buffer_vec.rs
@@ -103,6 +103,11 @@ impl<T: NoUninit> RawBufferVec<T> {
         self.values.append(&mut other.values);
     }
 
+    /// Returns the value at the given index.
+    pub fn get(&self, index: u32) -> Option<&T> {
+        self.values.get(index as usize)
+    }
+
     /// Sets the value at the given index.
     ///
     /// The index must be less than [`RawBufferVec::len`].
diff --git a/crates/bevy_render/src/view/mod.rs b/crates/bevy_render/src/view/mod.rs
index b0fab01d01b4e..ba0f8cca62ced 100644
--- a/crates/bevy_render/src/view/mod.rs
+++ b/crates/bevy_render/src/view/mod.rs
@@ -2,6 +2,7 @@ pub mod visibility;
 pub mod window;
 
 use bevy_asset::{load_internal_asset, weak_handle, Handle};
+use bevy_diagnostic::FrameCount;
 pub use visibility::*;
 pub use window::*;
 
@@ -568,6 +569,7 @@ pub struct ViewUniform {
     pub frustum: [Vec4; 6],
     pub color_grading: ColorGradingUniform,
     pub mip_bias: f32,
+    pub frame_count: u32,
 }
 
 #[derive(Resource)]
@@ -889,6 +891,7 @@ pub fn prepare_view_uniforms(
         Option<&TemporalJitter>,
         Option<&MipBias>,
     )>,
+    frame_count: Res<FrameCount>,
 ) {
     let view_iter = views.iter();
     let view_count = view_iter.len();
@@ -942,6 +945,7 @@ pub fn prepare_view_uniforms(
                 frustum,
                 color_grading: extracted_view.color_grading.clone().into(),
                 mip_bias: mip_bias.unwrap_or(&MipBias(0.0)).0,
+                frame_count: frame_count.0,
             }),
         };
 
diff --git a/crates/bevy_render/src/view/view.wgsl b/crates/bevy_render/src/view/view.wgsl
index ed08599758a07..317de2eb88073 100644
--- a/crates/bevy_render/src/view/view.wgsl
+++ b/crates/bevy_render/src/view/view.wgsl
@@ -60,4 +60,5 @@ struct View {
     frustum: array<vec4<f32>, 6>,
     color_grading: ColorGrading,
     mip_bias: f32,
+    frame_count: u32,
 };